1993-1999 Master studies in Pharmaceutical Chemistry at the Universita’ degli Studi di Trieste, Trieste, Italia (110/110).
1997-1999 Master Thesis at the Faculty of Medicine of the Universita’ degli Studi di Trieste, Trieste, Italia. “TNF-Induced Shedding of TNF Receptors in Human Neutrophils: Role of the 55-kDa TNF Receptor and Involvement of a Membrane-Bound and Non-Matrix Metalloproteinase”
2000-2004 PhD in Neuroscience at the University of Heidelberg, Germany.
“Engrailed Genes are cell autonomously required for the survival of mesencephalic dopaminergic neurons”.
2013-2015 Habilitation in Neuroanatomy at the University of Fribourg “Cellular Signaling in mature brain function and dysfunction”
2016-2018 Master in Business Administration, University of Illinois, Urbana Champaign, Illinois, US.